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Mrs. Jeffries is the ex-wife of Professor Proton and mother of their son.

It's revealed in "The Proton Resurgence" that the puppeteer who did Gino the Neutrino also did Mrs. Jeffries (made a move on her) and this caused Arthur to hate the puppet Gino.

In "The Proton Transmogrification", Sheldon asked Arthur Jeffries's spirit why he is in Apartment 4A and Arthur replies that he doesn't know. He adds that maybe he came back to haunt his ex-wife.

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